I don't think types can be considered masculine or feminine really, but if there is anything, probably Beta and Gamma are more masculine and Alpha and Delta are more feminine because of the Se in the former quadras and Si in the latter. Even that seems to be a quite weak correlation. I also think that, regarding judging functions, Fe and Ti are more masculine, while Fi and Te are more feminine (mostly just Fi). I mean, Fe includes all the traditionally "macho" kinds of emotions (i.e. rage, standing up for your externally-based values like a heroic warrior), which are also valued in the Romance cultures that originated the concept of "macho", although of course the Tx will probably come across more masculine generally.

You also know a paper is bad when it mentions unfalsifiable evolutionary psychology BS from the outset.